Recent Publications (this century)


Kustka, A.B., A.E. Allen and F.M.M. Morel, Sequence analysis and transcriptional regulation of Fe acquisition genes in two marine diatoms. Journal of Phycology. (in press)

Xu, Y., D. Tang, Y.Shaked and F.M.M. Morel, Zinc, cadmium and cobalt inter-replacement and relative use efficiencies in the coccolithophore Emiliania huxleyi.,Limnology and Oceanography (in press)

Park, H., B. Song, and F.M.M. Morel, Diversity of the cadmium containing carbonic anhydrase (DDCA) in marine diatoms and seawater., Environmental Microbiology9 (2) p. 403-413 doi:10.1111/j.1462-2920.2006.01151.x (2007).


Castruita, M., M. Saito, P.C. Schottel, L.A.Elmegreen, S. Myneni, E.I. Stiefel, and F.M.M. Morel, Overexpression and characterization of an iron storage and DNA-binding Dps protein from Trichodesmium erythraeum. Applied Env. Microbiology, 2918-2924 (2006).

Shaked, Y., Y. Xu, T.-Y.-Ho, K. Leblanc and F.M.M. Morel,  Zinc availability and Alkaline phosphatase activity in Emiliania huxleyi: Implications for Zn-P co-limitation in the ocean,  Limnology and Oceanography, 51; 299-309 (2006).

Tang, D. and F.M.M. Morel, Distinguishing between intracellular and extracellular trace metals in phytoplankton. Marine Chemistry, 8 p. 18-30 (2006).

Xu, Y, T.M. Wahlund, L. Feng, Y. Shaked, and F.M.M.Morel, A novel alkaline phosphatase in the coccolithophore emiliania huxleyi (prymnesiophyceae) and its regulation by phosphorus. Journal of Phycology. Vol.42(4); 835-844 (2006).


Amyot, M., F.M.M. Morel and P. A. Ariya, Dark oxidation of dissolved and liquid elemental mercury in aquatic environments. Environ. Sci. Technol. 39(1) 110-114, (2005).

Kustka, A. B., Y. Shaked, A.J. Milligan, D. W. King, F.M.M. Morel,  Extracellular production of superoxide by marine diatoms: contrasting effects on iron redox chemistry and bioavailabilty. Limnol. and Oceanogr.,  50(4) 1172-1180 (2005).

Lane, T., M.A. Saito, H.Park, G.N. George, I.J. Pickering, R.C. Prince, F.M.M. Morel, A cadmium enzyme from a marine diatom. Nature Vol. 435, p. 42 (2005)

Morel,  F.M.M.  and E. G. Malcolm, “The Biogeochemistry of Cadmium” in Biogeochemical Cycles of Elements, Vol. 43 of  ‘Metal Ions in Biological Systems’; A. Sigel, H. Sigel, and R. K. O. Sigel, Eds.; M. Dekker, New York, pp. 301 (2005).

Shaked, Y., A.B. Kustka and F.M.M. Morel, A general kinetic model for iron acquisition by eukaryotic phytoplankton. Limnol. and Oceanogr., 50(3) 872-882 (2005).

Sternberg, E., D. Tang, T.-Y. Ho, C. Jeandel and F.M.M. Morel,   Barium Uptake and Adsorption in Diatoms, Geochemica Acta , Vol. 69(11) 2745-2752 (2005).

Sunda, W., N.M. Price, F.M.M. Morel, “Trace Metal ion buffers and their use in culture studies” in Algal Culturing Techniques, R. Anderson, Editor, Academic Press, Burlington, MA p. 35-63 (2005).


Kraepiel, A.M.L., K. Keller, H.B. Chin, E.G. Malcolm, F.M.M. Morel,    Response to Comment on  “Sources and Variations of Mercury in Tuna” Environ. Sci. Technol., 38(14) 4048 (2004).

Lalonde, J.D., M. Amyot, J. Orvoine, F.M.M. Morel, J-C. Auclair, P.A. Ariya,   Photoinduced Oxidation of Hg0(aq) in the Waters from the St. Lawrence Estuary.,  Environ. Sci. Technol., 38 (2) 508-514. (2004).

Milligan, A. J. and F.M.M. Morel, Dynamics of silicon metabolism and silicon isotopic discrimination in a marine diatom as a function of pCO2Limnol. Oceanogr., 49 (2), 322-329 (2004).

Reinfelder, J.R., A.J. Milligan, F.M.M. Morel ,  The role of the C4 pathway in carbon accumulation and fixation in a marine diatom.  Plant Physiology Vol. 135, 2106-2111 (2004).

Shaked, Y., A.B. Kustka, F.M.M. Morel and Y. Erel,  Simultaneous determination of iron reduction and uptake by phytoplankton, Limnol.  and Oceanography: Methods,  Vol . 2, 137-145 (2004).


Ekstrom, E.B., F.M.M. Morel, J.M. Benoit,   Mercury Methylation Independent of Acetyl-CoA Pathway in SRB. Appl. and Environ. Microbiol.,  69 (9) 5414 – 5422 (2003).

Ho, T.-Y., A. Quigg, Z.V. Finkel, A.J. Milligan, K. Wyman, P.G. Falkowski, F.M.M. Morel,  The Elemental Composition of Some Marine Phytoplankton.  J. Phycol., 39, 1145-1159. (2003).

Kraepiel, A.M.L., K. Keller, H.B. Chin, E.G. Malcolm, F.M.M. Morel,   Sources and Variations of Mercury in Tuna.  Environ. Sci. Technol., 37 (24)5551-5558 (2003).

Quigg, A., Z.V. Finkel, A.J. Irwin, Y. Rosenthal, T.-Y. Ho, J.R. Reinfelder, O. Schofield, F.M.M. Morel, P.G. Falkowski,  The evolutionary inheritance of elemental stoichiometry in marine phytoplankton.  Nature, 425: 291-294. (2003).

Morel, F.M.M. and N.M. Price, The Biogeochemical Cycles of Trace Metals in the Oceans.  Science, 300: 944-947 (2003).

Morel, F.M.M., A.J. Milligan and M.A. Saito, “Marine Bioinorganic Chemistry:  The Role of Trace of Metals in the Oceanic Cycles of Major Nutrients” in Treatise on Geochemistry, Vol. 6, edited by KK. Turekian, H.D. Holland, Elsevier Science Ltd, Cambridge, UK, p. 113-143 (2003).

Saito, M. A., D.M. Sigman and F.M.M. Morel,  The bioinorganic chemistry of the ancient ocean:  the co-evolution of cyanobacterial metal requirements and biogeochemical cycles at the Archean – Proterozoic boundary?  Inorganica Chimica Acta  356C, 308-318 (2003).


Jay, J.A., K.J. Murray, C.C. Gilmour, R.P. Mason, F.M.M. Morel, A.L. Roberts,  and H.F. Hemond, Mercury methylation by Desulfovibrio desulfuricans ND132 in the presence of polysulfides. Appl. & Env. Microbiol., 68(11): 5741-5745 (2002).

Milligan, A.J. and F.M.M. Morel,  A proton buffering role for silica in diatoms.  Science, 297: 1848-1850 (2002).

Morel, F.M.M., E.H. Cox, A.M.L. Kraepiel, T.W. Lane, A.J. Milligan, I. Schaperdoth, J.R. Reinfelder and P.D. Tortell,  Acquisition of inorganic carbon by the marine diatom Thalassiosira weissflogiiFunctional Plant Biology, 29: 2,3 p. 301-308 (2002).

Tortell, P.D., G.R. DiTullio, D.M. Sigman, and F.M.M. Morel,  CO2 effects on taxonomic composition and nutrient utilization in an Equatorial Pacific phytoplankton assemblage.  MEPS, 236: 37-43 (2002).

Tortell, P.D. and F.M.M. Morel.  Sources of inorganic carbon for phytoplankton in the Eastern Subtropical and Equatorial Pacific Ocean.  Limnol. Oceanogr., 47: 1012-1022 (2002).


Gawel, J.E., C.G. Trick, and F.M.M. Morel,  Phytochelatins are bioindicators of atmospheric metal insult to trees near Sudbury, Ontario, Canada.  Environ. Sci. Technol., 35: 2108-2113 (2001).

Lam, P.J., P.D. Tortell, and F.M.M. Morel,  Differential effects of iron additions on organic and inorganic carbon production by phytoplankton.  Limnol. Oceanogr., 46: 1199-1202 (2001).

Lalonde, J.D., M. Amyot, A.M.L. Kraepiel, and F.M.M. Morel,  Photooxidation of Hg(0) in an estuarine system.  Environ. Sci. Technol., 35: 1367-1372 (2001).


Cox, E.H., G.L. McLendon, F.M.M. Morel, T.W. Lane, R.C. Prince, I.J. Pickering, and G.N. George, The active site structure of Thalassiosiraweissflogii carbonic anhydrase 1.  Biochemistry, 39: 12128-12130 (2000).

Jay, J., F.M.M. Morel, and H.F. Hemond, Mercury speciation in the presence of polysulfides. Environ. Sci. Technol., 34: 2196-2200 (2000).

Keller, K., K. Tan, F.M.M. Morel, and D.F. Bradford, Preserving the ocean circulation: Implications for climate policy. Climatic Change 47(1-2): 17-43 (2000).

Lane, T.W. and F.M.M. Morel.  Regulation of carbonic anhydrase expression by zinc, cobalt, and carbon dioxide in the marine diatom Thalassiosira weissflogiiPlant Physiology, 123: 345-352 (2000).

Lane, T.W. and F.M.M. Morel.  A biological function for cadmium in marine diatoms.  PNAS, 97: 4627-4631 (2000).

  Morel, F.M.M., Broadening the Scientific Ph.D.:  The Princeton Experience.  Proceedings of NRC’s Chemical Sciences Roundtable (CSR) Workshop, Graduate Education in the Chemical Sciences:  Issues for the 21st Century, p. 130-134 (2000). 

Morel, F.M.M., Use of Trace Metals in Marine Bioremediation:  A Need for Fundamental Knowledge.  Reviews of Proceedings of the Workshop on Opportunities for Environmental Applications of Marine Biotechnology, p. 184-192 (2000). 

Reinfelder, J.R., A.M.L. Kraepiel, and F.M.M. Morel.  Unicellular C4 photosynthesis in a marine diatom.  Nature, 407: 996-999 (2000).

Riebesell, U., I. Zondervan, B. Rost, P.D. Tortell, R.E. Zeebe, and F.M.M. Morel,  Reduced calcification in marine plankton in response to increased atmospheric CO2.  Nature, 407: 364-367 (2000).

Tortell, P.D., G.H. Rau, and F.M.M. Morel,  Inorganic carbon acquisition in coastal Pacific phytoplankton communities.  Limnol. Oceanogr., 45: 1485-1500 (2000).

Past Publications (last century)


Ahner, B. and F.M.M. Morel.  Phytochelatins in microalgae.  Progress in Phycological

Research, 13: 1-31 (1999).

Cullen, J.T., T.W. Lane, F.M.M. Morel, and R.M. Sherrell.  Modulation of cadmium uptake in phytoplankton by seawater CO2 concentration.  Nature, 402: 165-167 (1999).

Keller, K. and F.M.M. Morel.  A model of carbon isotopic fractionation and active carbon uptake in phytoplankton.  Marine Ecology Progress Series, 182: 295-298 (1999).

Kraepiel, A.M.L., K. Keller, and F.M.M. Morel.  A model for metal adsorption on montmorillonite. J.Colloid and Interface Science, 210: 43-54 (1999).


Ahner, B.A., J.G. Lee, N.M. Price, and F.M.M. Morel.  Phytochelatin concentrations in the equatorial Pacific.  Deep-Sea Res., 45:1779-1796 (1998).

Gueneau, P., F.M.M. Morel, J. Laroche, and D. Erdner.  The PetF region of the chloroplast genome from the diatom Thalassiosiraweissflogii.: sequence, organization and phylogeny.  European Journal of Phycology, 33: 203-211 (1998).

Kraepiel, A.M.L., K. Keller, and F.M.M. Morel.  On the acid-base chemistry of permanently charged minerals.  Envi. Sci. Tech., 32: 2829-2838 (1998).

Morel, F.M.M., A.M.L. Kraepiel, and M. Amyot.  The chemical cycle and bioaccumulation of mercury.  Annual Review Ecology and Systematics, Vol. 29: 543-566 (1998).

Newman, D.K., D. Ahmann, and F.M.M. Morel.  A brief review of dissimilatory arsenate

reduction.  Geomicrobio. Jour., 15: 255-268 (1998).

Price, N.M. and F.M.M. Morel,  Biological cycling of iron in the ocean.  In:  Iron Transport and Storage in Microorganisms, Plants, and Animals, Vol. 35, p. 1-36 of 'Metal Ions in Biological Systems', A. Sigel and H. Sigel, eds.; M. Dekker, Inc., New York, (1998).


Ahmann, D., L.R. Krumholz, H.H. Hemond, D.R. Lovley, and F.M.M. Morel.  Microbial mobilization of arsenic from sediments of the Aberjona Watershed.  Env. Sci. Tech., 31: 2923-2930 (1997).

Ahner, B.A., F.M.M. Morel, and J.W. Moffett.  Trace metal control of phytochelatin production in coastal waters.  Limnol. Oceanogr., 42: 601-608 (1997).

Amyot, M., G.A. Gill, and F.M.M. Morel.  Production and loss of dissolved gaseous mercury in coastal seawater.  Env. Sci. Tech., 31: 3606-3611 (1997).

Kraepiel, A.M.L., J.-F. Chiffoleau, J.-M. Martin, and F.M.M. Morel.  Geochemistry of trace metals in the Gironde estuary.  Geochim. Cosmochim. Acta, 61: 1421-1435 (1997).

Morel, F.M.M. and A. Kraepiel.  Coulombic effects on the adsorption of trace cations on clays.  J. Cont. Hydrology,  28: 17-20 (1997).

Morel, F.M.M.  Discussion on “A mechanistic description of Ni and Zn sorption on Na-montmorillonite. Part I: Titration and sorption measurements.  Part II: Modeling” by Bart Baeyens and Michael H. Bradbury.  J. Cont. Hydrology,  28: 7-10 (1997).

Newman, D.K., E.K. Kennedy, J.D. Coates, D. Ahmann, D.J. Ellis, D.R. Lovley, and F.M.M. Morel.  Dissimilatory As(V) and S(VI) reduction in Desulfotomaculum aurpigmentum, sp.novArchives Microbiol., 168: 380-388 (1997).

Newman, D.K., T.J. Beveridge and F.M.M. Morel.  Precipitation of As2S3 by a novel As(V)-reducing bacterium, strain OREX-4.  Appl. Env. Microbiol., 63: 2022-2028 (1997).

Roberts, S.B., T. Lane, and F.M.M. Morel.  Carbonic anhydrase in the marine diatom Thalassiosira weissflogii (bacillariophyta).  J. Phycol., 33: 845-850 (1997).

Tortell, P.D., J.R. Reinfelder, and F.M.M. Morel.  Active uptake of bicarbonate by diatoms. Nature, 390: 243-244 (1997).

Vöelker, B., F.M.M. Morel and B. Sulzberger.  Iron redox cycling in surface waters: effects of humic substances and light.  Env. Sci. Tech., 31: 1004-1011 (1997).


Gawel, J.E., B.A. Ahner, A.J. Friedland and F.M.M. Morel.  Role for heavy metals in forest decline indicated by phytochelatin measurements.  Nature , 381: 64-65 (1996).

Lee, J.G., B.A. Ahner and F.M.M. Morel.  Export of cadmium and phytochelatin by the marine diatom  Thalassiosira weissflogii.  Env. Sci. Tech., 30: 1814-1821 (1996).

Mason, R.P., J.R. Reinfelder and F.M.M. Morel.  The uptake, toxicity, and trophic transfer of mercury in a coastal diatom.  Env. Sci. Tech., 30: 1835-1845 (1996).

Yee, D. and F.M.M. Morel.  In vivo substitution of zinc by cobalt in carbonic anhydrase of a marine diatom.  Limnol. Oceanogr., 41: 573-577 (1996).


Ahner, B.A., S. Kong and F.M.M. Morel.  Phytochelatin synthesis in marine algae: I. An interspecies comparison.  Limnol. & Oceanography, 40(4): 649-657 (1995).

Ahner, B.A. and F.M.M. Morel.  Phytochelatin production in marine algae: II. Induction by various metals.  Limnol. & Oceanography, 40(4): 658-665 (1995).

Joseph, E.M. and F.M.M. Morel and N.M. Price.  Effect of aluminum on surface phosphatases of Chlamydomonas reinhardtiiCan. J. Fish. Aquat. Sci., 52: 353-357 (1995).

Lee, J.G. and F.M.M.Morel.  Replacement of zinc by cadmium in marine phytoplankton.  Mar. Ecol. Prog. Ser., 127: 305-309 (1995).

Lee, J.G., S.R. Roberts and F.M.M.Morel.  Cadmium: a nutrient for the marine diatom Thalassiosira weissflogiiLimnol. & Oceanogr., 40: 1056-1063 (1995).

Mason, R.P., F.M.M. Morel and H.F. Hemond.  The role of microorganisms in elemental mercury formation in natural waters.  Water Air Soil Pollut., 80: 775-787 (1995).

Mason, R.P., J.R. Reinfelder, and F.M.M.Morel.  Bioaccumulation of mercury and methylmercury.  Water Air Soil Pollut., 80: 915-921 (1995).

Morel, F.M.M. and J.R. Reinfelder.  Growth limits on phytoplankton – reply. Nature, 373 (6509): 28-28 (1995).


Ahmann, D., A.L. Roberts, L.R. Krumholz, and F.M.M. Morel. Microbe grows by reducing arsenic.  Nature,  371: 750 (1994).

Ahner, B.A., N.M. Price, and F.M.M. Morel.  Phytochelatin production by marine phytoplankton and low free metal ion concentrations: Lab studies and field data for Massachusetts Bay. Proc. Natl. Acad. Sci., 91: 8433-8436 (1994).

Mason, R.P., W.F. Fitzgerald and F.M.M. Morel.  The biogeochemical cycling of elemental mercury:  Anthropogenic Influences.  Geochim. Cosmochim. Acta, 58: 3191-3198 (1994).

Morel, F.M.M., J.R. Reinfelder, S.B. Roberts, C.P. Chamberlain, J.G. Lee, and D. Yee.  Zinc and carbon co-limitation of marine phytoplankton.  Nature,  369: 740-742 (1994).

Price, N.M., B.A. Ahner and F.M.M. Morel.  The Equatorial Pacific Ocean:  grazer-controlled phytoplankton populations in an iron-limited ecosystem.  Limnol. & Oceanogr., 39: 520-534 (1994).


Hudson, R.J.M. and F.M.M. Morel.  Trace metal transport by marine microorganisms:  implications of metal coordination kinetics.  Deep-Sea Res., 40: 129-150 (1993).

Mason, R.P., F.M.M. Morel, and H.F. Hemond.  The role of microorganisms in elemental mercury formation in natural waters. In:  Heavy Metals in the Environment (J.O. Nriagu and R.P. Allan, eds.) Vol. 2 pp. 293-296 (1993).

Morel, F.M.M. and J.G. Hering,  Principles and Applications of Aquatic Chemistry (2nd ed.).  John Wiley, New York, (1993).


Bartschat, B.M., S.E. Cabaniss and F.M.M. Morel.  An oligoelectrolyte model for cation binding by humic substances.  Env. Sci. Tech., 26: 284-294 (1992).

Green, S.A., F.M.M. Morel and N.V. Blough.  Investigation of the electrostatic properties of humic substances by fluorescence quenching.  Env. Sci. Tech., 26: 294-302 (1992).

Hudson, R.J.M., D.T. Covault and F.M.M. Morel.  Investigations of iron coordination and redox reactions in seawater using 59Fe radiometry and ion-pair solvent extraction of amphiphilic iron complexes.  Marine Chem., 38: 209-235 (1992).


Chisholm, S.W. and F.M.M. Morel (eds.) What regulates phytoplankton production in nutrient-rich areas of the open sea?  Limnol. & Oceanogr., Vol. 36, No. 8, Dec. (1991).

Chisholm, S.W., F.M.M. Morel. What controls phytoplankton production in nutrient-rich areas of the open sea. Limnol. & Oceanogr., 36(8): U1507-U1511 (1991).

Morel, F.M.M., R.J.M. Hudson and N.M. Price.  Limitation of productivity by trace metals in the sea.  Limnol. & Oceanogr., 36: 1742-1755 (1991).

Morel, F.M.M.,  J.G. Rueter, and N.M. Price.  Iron nutrition of phytoplankton and its possible importance in the ecology of ocean regions with high nutrient and low biomass. Oceanography, 4: 56-61 (1991).

Morel, F.M.M.,  D.A. Dzombak and N.M. Price, Heterogeneous reactions in coastal waters.  In:  Ocean Margin Processes in Global Change, Mantoura, Martin, and Wollast (eds.), Dahlem Konferenzen.  John Wiley, New York (1991).

Palenik, B., D.J. Kieber and F.M.M. Morel.  Dissolved organic nitrogen use by phytoplankton:  The role of cell-surface enzymes.  Biological Oceanography, 6: 347-354 (1991).

Palenik, B. and F.M.M. Morel.  Amine oxidases of marine phytoplankton.  Appl. Env. Microbiol., 57: 2440-2443 (1991).

Palenik, B., N.M. Price and F.M.M. Morel.  Potential effects of UV-B on the chemical environment of marine organisms: A review.  Environmen. Pollut., 70: 117-130 (1991).

Price, N.M. and F.M.M. Morel.  Co-limitation of phytoplankton growth by nickel and nitrogen. Limnol. & Oceanogr., 36: 1071-1077 (1991).

Price, N.M., L.F. Andersen and F.M.M. Morel.  Iron and nitrogen nutrition of equatorial Pacific plankton.  Deep-Sea Res., 38: 1361-1378 (1991).


Dzombak, D.A. and F.M.M. Morel.  Surface Complexation Modeling:  Hydrous Ferric Oxide.  John Wiley, New York (1990).

Hering, J. and F.M.M. Morel, The kinetics of trace metal complexation:  Implications for metal reactivity in natural waters.  In:  Aquatic Chemical Kinetics, W. Stumm, (ed.), John Wiley, New York (1990).

Hering, J.G. and F.M.M. Morel.  Kinetics of trace metal complexation: Ligand exchange reactions.  Env. Sci. Tech., 24: 242-252 (1990).

Hudson, R.J.M. and F.M.M. Morel.  Iron transport in marine phytoplankton:  Kinetics of cellular and medium coordination reactions.  Limnol. & Oceanogr., 35: 1002-1016 (1990).

Newman, K.A., F.M.M. Morel and K.D. Stolzenbach.  Settling and coagulation characteristics of fluorescent particles determined by flow cytometry and fluorometry.  Env. Sci. Tech.,  24: 506-513 (1990).

Nirel, P.M. and F.M.M. Morel.  Comment: Pitfalls of sequential extractions.  Water Research,  24: 1055-1056 (1990).

Palenik, F. and F.M.M. Morel.  Amino acid utilization by marine phytoplankton:  A novel mechanism.  Limnol. & Oceanogr., 35: 260-269 (1990).

Palenik, B. and F.M.M. Morel.  Comparison of cell-surface L-amino acid oxidases from several marine phytoplankton.  Mar. Ecol. Prog. Ser., 59: 195-201 (1990).

Price, N.M. and F.M.M. Morel, The role of extracellular enzymatic reactions in natural waters.  In:  Aquatic Chemical Kinetics, W. Stumm, (ed.), John Wiley, New York, Chapter 8, 235-257 (1990).

Price, N.M. and F.M.M. Morel.  Cadmium and cobalt substitution for zinc in a zinc-deficient marine diatom.  Nature, 344: 658-660 (1990).

Rich, H.W. and F.M.M. Morel.  Availability of well-defined iron colloids to the marine diatom Thalassiosira weissflogii.Limnol. & Oceanogr., 35: 652-662 (1990).

Roulier, M.A. B. Palenik and F.M.M. Morel.  A method for the measurement  of choline and hydrogen peroxide in seawater.  Marine Chem., 30: 409-421 (1990).


Cabaniss, S.E. and F.M.M. Morel.  Comment on “A unified physicochemical description of the protonation and metal ion complexation equilibrium of natural organic acids, Part 1 & 2.” Env. Sci. Tech., 23: 746-747 (1989).

Hering, J.G. and F.M.M. Morel.  Slow coordination reactions in seawater. Geochim. Cosmochim. Acta, 53: 611-618 (1989).

Hudson, R.J.M. and F.M.M. Morel.  Distinguishing between extra- and intracellular iron in marine phytoplankton.  Limnol. & Oceanogr., 34: 1113-1120 (1989).

Morel, F.M.M. and B. Palenik, The aquatic chemistry of trace metals in biofilms.  In:  Structure and Function of Biofilms, W.G. Characklis and P.A. Wilderer (eds.), Dahlem Konferenzen.  John Wiley, New York (1989).

Price, N.M., G.I. Harrison, J.G. Hering, R.J. Hudson, P.M.V. Nirel, B. Palenik and F.M.M. Morel.  Preparation and chemistry of the artificial algal culture medium aquil.  Biolog.Oceanogr., 6: 443-461 (1988/1989).


Hering, J.G. and F.M.M. Morel.  Kinetics of trace metal complexation:  Role of alkaline earth metals.  Env. Sci. Tech., 22: 1469-1478 (1988).

Hering, J.G. and F.M.M. Morel.  Humic acid complexation of calcium and copper.  Env.Sci. Tech., 22: 1234-1237 (1988).

Jones, G.J. and F.M.M. Morel.  Plasmalemma redox activity in the diatom ThalassiosiraPlant Physiol., 87: 143-147 (1988).

Morel, F.M.M.  Solute-particle and particle-particle reactions in seawater.  Appl. Geochem. 3:80 (1988).

Palenik, B. and F.M.M. Morel.  Dark production of H2O2 in the Sargasso Sea.  Limnol. & Oceanogr., 33(6:2) 1606-1611 (1988).


Coudrain-Ribstein, A. and F.M.M. Morel.  Transport modeling of major reactive elements in a variable temperature field:  Methodology and simple examples.  Bulletin de la Societe Geologique de France Numero Special 5: 1009-1017 (1987).

Dzombak, D.A. and F.M.M. Morel.  Adsorption of inorganic pollutants in aquatic systems.  J. Hydraulic Eng. ASCE. 113: 430-475 (1987).

Dzombak, D.A. and F.M.M. Morel.  Development of a data base for modelling adsorption of inorganics on iron and aluminum oxides.  Environmental Progress  AICHE 6: 133-137 (1987).

Hering, J.G., , W.G. Sunda , R.L. Ferguson  and F.M.M. Morel.  A field comparison of two methods for the determination of copper complexation:  Bacterial bioassay and fixed-potential amperometry.  Marine Chemistry, 20: 255-312 (1987).

Jones, G., F.M.M. Morel, C. Yentsch and E.A. Funkhouser, A possible immunological approach to the study of plasmalemma redox enzymes in phytoplankton.  Lecture series in coastal and etuarine studies.  Springer-Verlag (1987).

Jones, G.J., B. Palenik and F.M.M. Morel.  Trace metal reduction by phytoplankton:  The role of plasmalemma redox enzymes.  J. Phycol., 23: 237-244 (1987).

Morel, F.M.M. and P.M. Gschwend, The role of colloids in the partitioning of solutes in natural waters.  In:  Aquatic Surface Chemistry, W. Stumm (ed.), John Wiley, New York (1987).

Morel, F.M.M.  Kinetics  of nutrient uptake and growth in phytoplankton.  J. Phycol., 23: 137-150 (1987).

Palenik, B., F.M.M. Morel and O.C. Zafiriou.  Hydrogen peroxide production by a marine phytoplankton.  Limnol. & Oceanogr., 32(6): 1365-1369 (1987).


Cherry, J.A., F.M.M. Morel, J.V. Rouse, J.L. Schnoor and M.G. Wolman.  Hydrogeochemistry of sulfide and arsenic-rich tailings and alluvium along Whitewood Creek, South Dakota.  Three part series.  Mineral and Energy Resources, Colorado School of Mines (1986).

Dzombak, D.A. and F.M.M. Morel.  Sorption of cadmium on hydrous ferric oxide at high sorbate/sorbent ratios:  Equilibrium kinetics and modeling.  J. Coll. & Inter. Sci., 112(2): 588-598 (1986).

Dzombak, D.A., W. Fish and F.M.M. Morel.  Metal-humate interactions, 1:  Discrete ligand and continuous distribution models.  Env. Sci. Tech., 20(7): 669-675 (1986).

Farley, K. and F.M.M. Morel.  The role of coagulation in the kinetics of sedimentation.  Env. Sci. Tech., 20(2): 187-195 (1986).

Fish, W., D.A. Dzombak and F.M.M. Morel.  Metal-humate interactions, 2:  Applications and comparison of models.  Env. Sci. Tech., 20(7): 676-683 (1986).

Harrison, G.I. and F.M.M. Morel.  Response of the marine diatom Thalassiosira weissflogii to iron stress.  Limnol. & Oceanogr., 31(5): 989-997 (1986).

Morel, F.M.M., Trace metals-phytoplankton interaction:  An overview.  In:  Biogeochemical Processes at the Land Sea Boundary, P. Lassere and J.M. Martin (eds.).  Elsevier, Amsterdam (1986).


Farley, K.J., D.A. Dzombak and F.M.M. Morel.  A surface precipitation model for the sorption of cations on metal oxides.  J. Coll. Inter. Sci., 106:226 (1985).

Fish, W. and F.M.M. Morel.  Propagation of error in fulvic acid titration data:  A comparison of three analytical methods.  Can. J. Chem., 63(6): 1185-1193 (1985).

Morel, F.M.M. and R.J.M. Hudson,  The geobiological cycle of trace elements in aquatic systems:  Redfield revisited.  In:  Chemical Processes in Lakes, W.Stumm (ed.).  John Wiley, New York (1985).

Watras, C.J., MacFarlane J. and F.M.M. Morel.  Nickel accumulation by Scenedesmus  and Daphnia:  Food-chain transport and geochemical implications.  Can. J. Fish. Aquat. Sci., 42(4): 724-730 (1985).


Morel, F.M.M. and S.L. Schiff,  Geochemistry of municipal waste in coastal waters.  In:  Ocean Discharge of Municipal Wastes:  Impact on Coastal Waters, E.P. Myers (ed.), MIT Sea Grant College Program Publishers (1984).

Waite, T.D. and F.M.M. Morel.  Photoreductive dissolution of colloidal iron oxides in natural waters.  Env.Sci. Tech., 18: 860-868 (1984).

Waite, T.D. and F.M.M. Morel.  Ligand exchange and fluorescence quenching studies of the fulvic acid-iron interaction:  Effects of pH and light.  Anal. Chemica Acta, 162: 263-264 (1984).

Waite, T.D. and F.M.M. Morel.  Photoreductive dissolution of colloidal iron oxide:  Effect of citrate.  J. Coll. Inter. Sci., 102:121 (1984).

Waite, T.D. and F.M.M. Morel.  Coulometric technique for studying the redox dynamics of iron in seawater.  Anal. Chem., 56: 787-792 (1984).


Fish, W. and F.M.M. Morel.  Characterization of organic copper-complexing agents released by Daphnia magna.  Can. J. Fish. Aquatic Sci., 40:1270 (1983).

Harrison, G.I. and F.M.M. Morel.  Antagonism between cadmium and iron in the marine diatom Thalassiosira weissflogii.  J. Phycol., 19: 495-507 (1983).

Morel, F.M.M. and N.M.L. Morel-Laurens,  Trace Metals and plankton in the oceans:  Facts and speculations.  In:  Trace Metals in Sea Water, C.S. Wong, E. Boyle, K.W. Bruland, J.D. Burton and E.D. Goldberg (eds.), NATO Conference Series, Series IV:  Marine Chemistry, Plenum Press (pp. 841-869) (1983).

Morel, F.M.M., Principles of Aquatic Chemistry, John Wiley & Sons, Inc., New York, (1983).

Morel, F.M.M. and N.M.L. Morel-Laurens,  Trace Metals and plankton in the oceans:  Facts and speculations.  In:  Trace Metals in Sea Water, C.S. Wong, E. Boyle, K.W. Bruland, J.D. Burton and E.D. Goldberg (eds.), NATO Conference Series, Series IV:  Marine Chemistry, Plenum Press (pp. 841-869) (1983).

Waite, T.D. and F.M.M. Morel.  Characterization of complexing agents in natural waters by Cu(II)/Cu(I) amperometry.  Anal. Chem., 55: 1268-1274 (1983).


Anderson, M.A. and F.M.M. Morel.  The influence of aqueous iron chemistry on the uptake of iron by the coastal diatom Thalassiosira weissflogiiLimnol. & Oceanogr., 27: 789 (1982).

Foster, P.L. and F.M.M. Morel.  Reversal of cadmium toxicity in the diatom Thalassiosira weissflogii.  Limnol. & Oceanogr., 27: 745-752 (1982).


Rueter, J.G., Jr. and F.M.M. Morel.  Interactions between zinc deficiency and copper toxicity as it affects the silicic acid uptake in the marine diatom, Thalassiosira pseudonana.  Limnol. & Oceanogr., 26:67-73 (1981).

Rueter, J.G., Jr., S.W. Chisholm and F.M.M. Morel.  The effect of copper toxicity on silicic acid uptake and growth in Thalassiosira pseudonana (Bacillariophyceae).  J. Phycol., 17: 270-278 (1981).


Anderson, M.A. and F.M.M. Morel.  Uptake of Fe(II) by a diatom in oxic culture medium.  Mar. Bio. Let., 1:263 (1980).

Chisholm, S.W., F.M.M. Morel and W.S. Slocum, The phasing and distribution of cell division cycles in marine diatoms.  In:  Primary Productivity in the Sea, P. Falkowski (ed.), Brookhaven Symposium in Biology No. 31, Plenum Press (1980).

McDuff, R.E. and F.M.M. Morel.  Geochemical control of seawater alkalinity (Sillén revisited).  Env. Sci. Tech., 14:1182 (1980).

McKnight, D.M. and F.M.M. Morel.  Copper complexation by siderophores from filamentous blue-green algae.  Limnol. & Oceanogr., 25:62 (1980).

Swallow, K.C., D.N. Hume and F.M.M. Morel.  Sorption of copper and lead by hydrous ferric oxide.  Env. Sci. Tech., 14:1326 (1980).


Anderson, D.M. and F.M.M.Morel,  Toxic dinoflagellate blooms in the Cape Cod region of Massachusetts.  In:  Toxic Dinoflagellate Blooms, D.L. Taylor and H.H. Seliger (eds.), Proceedings of the Second International Conference on Toxic Dinoflagellate Blooms, Key Biscayne, FL (1979).

Anderson, D.M. and F.M.M. Morel.  The seeding of two red tide blooms by the germination of benthic Gonyaulax tamarensis Hypnocysts.  J. Est. and Coast. Mar. Sci., 8:279 (1979).

McKnight, D.M. and F.M.M. Morel.  Release of weak and strong copper complexing agents by algae.  Limnol. & Oceanogr., 24(5):823-837 (1979).

Morel, F.M.M., N.M.L. Morel-Laurens, D.M. Anderson, D.M. McKnight and J.G. Rueter, Jr., Trace metal speciation and toxicity in phytoplankton cultures.  In:  Advances in Marine Environmental Research.  Proceedings of a Symposium, EPA-600/9-79-035 (1979).

Morel, F.M.M., J.G. Rueter, Jr., D.M. Anderson and R.R.L. Guillard.  AQUIL:  A chemical defined phytoplankton culture medium for trace metal studies.  J. Phycol., 15:135 (1979).

Westall, J.C., F.M.M. Morel and D.N. Hume.  Chloride interference in cupric ion selective electrode measurements.  Anal. Chem., 51:1792 (1979).


Anderson, M.A., F.M.M. Morel and R.R.L. Guillard.  Growth limitation of a coastal diatom by low zinc ion activity.  Nature, 276(5683):70-71 (1978).

Anderson, D.M. and F.M.M. Morel.  Copper sensitivity of Gonyaulax tamarensisLimnol. Oceanogr., 23:283 (1978).

Morel, N.M.L., J.G. Rueter, Jr. and F.M.M. Morel.  Copper toxicity to Skeletonema costatum (Bacillariophyceae).  J. Phycol., 14:43 (1978).

Swallow, K.C., D.M. McNight, J.C. Westall, N.M.L. Morel and F.M.M. Morel.  Potentiometric determination of copper complexation by phytoplankton exudates.  (Note).  Limnol. & Oceanogr., 23:538 (1978).

Yeasted, J.G. and F.M.M. Morel.  Empirical insights into lake response to nutrient loadings with application to models of phosphorus in lakes. Envir. Sci. Tech., 12:195 (1978).


Morel, F.M.M. and J.G. Yeasted, On the interfacing of chemical, physical and biological water quality models.  In:  Fate of Pollutants in the Air and Wave Environment, I.H. Suffet (ed.), Wiley-Interscience (1977).


Morel, F.M.M., R.E. McDuff and J.J. Morgan. Theory of interaction intensities, buffer capacities and pH stability in aqueous systems , with application to the pH of seawater and a heterogeneous model ocean system.  J. Mar. Chem., 4:1(1976).


Morel, F.M.M., J.C. Westall, C.R. O’Melia and J.J. Morgan.  Fate of trace metals in the Los Angeles County wastewater discharge. Env. Sci. Tech., 9:756 (1975).


Morel, F.M.M., R.E. McDuff and J.J. Morgan, Interactions and chemostasis in aquatic chemical systems.  In:  Trace Metals and Metal-Organic Interactions in Natural Waters, P.C. Singer (ed.), Ann Arbor Science, Ann Arbor, MI  (1973).


Morel, F.M.M.  A study of passive potassium efflux from human red blood cells using ion specific electrodes.  J. Membr. Bio., 12:68 (1972).

Morel, F.M.M. and J.J. Morgan. A numerical method for computing equilibria in aqueous chemical systems. Envir. Sci. and Tech.,  6:58 (1972).


Morel, F.M.M., R.F. Baker and H. Wayland.  Quantitation of human red blood cell fixation by glutaraldehyde.  J. Cell Biol., 48:91 (1971).

Principal Technical Reports and Technical Notes

TR 259  Morel, F.M.M. and S.L. Schiff.  Geochemistry of municipal waste in coastal waters.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA  (1980).

Contributing author to Dahlem Konferenzen report on “Organic Acids in Natural Waters,” John Wiley, New York (1989).

Contributing author to “Scientific Problems Relating to Ocean Pollution.” E.D. Goldberg (ed.), Proceedings of a Workshop, Estes Park, CO, July 1978, NOAA/ERL (1979).

Contributing author to “Trace Metals and Marine Production Processes,” R.A. Feely and H. Curl, Jr. (eds.), NOAA Special Report, NOAA/ERL #385 (1978).

TN 19  Westall, J.C. and F.M.M. Morel.  FITEQL:  A general algorithm for determination of metal-liqand complex stability constants from experimental data.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA  (1977).

TN 18  Westall, J.C., J.L. Zachary and F.M.M. Morel.  MINEQL:  A compact program for computation of chemical equilibria in aquatic systems.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA (1976).

TN 16  Morel, F.M.M., J.C. Westall, J.G. Rueter, Jr. and J.P. Chaplick.  Description of algal growth media AQUIL and FRAQUIL.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA (1975).

EQ. 73-02.  McDuff, R.E. and F.M.M. Morel.  Description of the chemical equilibrium program REDEQL2.  Keck Laboratory of Environmental Engineering Science, California Institute of Technology, Pasadena, CA (1973).

EQ. 72-01.  McDuff, R.E. and F.M.M. Morel.  REDEQL, a general program for the computation of chemical equilibrium in aqueous systems.  Keck Laboratory



Princeton University
Department of Geosciences