Morel Home

  François M. M. Morel
Professor of Geosciences (Geochemistry)

Department of Geosciences 
153 Guyot Hall 
Princeton University 
Princeton, NJ 08544

Phone: (609) 258-2416
Fax: (609) 258-5242

E-Mail: morel@princeton.edu

Morel Research Page

Curriculum Vitae

François M.M. Morel
Albert G. Blanke Professor of Geosciences
Princeton University

Director, Princeton Environmental Institute
Director, Center in Environmental BioInorganic Chemistry

Personal Information
Business: Department of Geosciences,
Guyot Hall, Room 153
Princeton University, Princeton NJ 08544
Telephone: (609) 258-2416
FAX: (609) 258-5242
E-mail: morel@princeton.edu


1966   Licence-ès-Sciences:  Mathématiques Appliquées.  Université de Grenoble
1967   Diplome d'Ingénieur.  Université de Grenoble, France
1971   Ph.D. Engineering Sciences.  California Institute of Technology, Pasadena
Academic Experience
1971-1973         Research Fellow in Environmental Engineering Sciences,
                          California Institute of Technology, Pasadena, CA

1973-1994         Assistant, Associate and then Professor, Department of Civil and
                          Environmental Engineering, Massachusetts Institute of Technology,
                          Cambridge, MA

1982-1995         Visiting Scientist, Woods Hole Oceanographic Institution, Woods Hole, MA

1986                  Associate Director for Research, Ecole des Mines de Paris, Paris, France

1987-1995         Visiting Professor, Ecole Normale Supèrieure, Paris, France

1991-1994         Director, R.M. Parsons Laboratory, Massachusetts Institute of Technology,
                          Cambridge, MA

1994-present     Professor, Department of Geosciences, Princeton University, NJ

1996-present     Visiting Professor, Université de Paris VI, France

1998-present     Director, Princeton Environmental Institute, Princeton University.

1998-present     Director, Center in Environmental BioInorganic Chemistry, Princeton University.

1999-present     Member of Editorial Board - Princeton University Press, Princeton University.

2001                  Visiting Professor Université Louis Pasteur, Strasbourg, France.


Fulbright International Travel Scholarship (1967)
French Government Ministry of Foreign Affairs Scholarship (1967)
Doherty Professorship in Ocean Utilization (1974-1977)
Chairman, Gordon Research Conference on Environmental Sciences (1986)
Association of Environmental Engineering Professors.  Ph.D. Thesis Award/as advisor (1988)
Turner Professorship in Civil & Environmental Engineering (1993)
AEEP Outstanding Paper Award (1994)
Albert G. Blanke Professorship (1996)
Chairman, Gordon Research Conference on Chemical Oceanography (1998)
Fellow of the Geochemistry Society (2000)
C.C. Patterson Medal, Geochemical Society (2001)
Guggenheim Fellowship (2001-2002)
Chairman, Gordon Research Conference on Environmental BioInorganic Studies (2002)

Professional Activities

Teaching:  Instructor of undergraduate and graduate subjects in Aquatic Chemistry Element Cycles; Surface Phenomena; Biogeochemistry.

Research:  Principal Investigator of projects on: (i) Coordination chemistry, surface chemistry and photochemistry of trace elements in natural waters; (ii) Interactions between the chemistry and microbiota in aquatic systems, (iii) Trace metal nutrition and toxicity in phytoplankton; and (iv) Environmental Bio-Inorganic Chemistry.

Visiting Committee's:  Swiss Federal Institute for Environmental Science and Technology, (EAWAG) 1994; Swiss Federal Institute of Technology, (Lausane) 1996; California Institute of Technology, (Caltech) Division of Geological and Planetary Science, 2000.

Society Memberships and Activities

American Society for Microbiology

American Chemical Society

Advisory Board of Environmental Science and Technology, 1978-1983; Associate Editor 1980

American Society of Limnology and Oceanography

Editorial Board of Limnology and Oceanography, 1980-1983, Editor, Special 1990

The Geochemistry Society

New England Aquarium, Member, Scientific Council, 1988-1990

Association of Environmental Engineering Professors

American Geophysical Union

Editorial Board, Biogeochemistry

Editorial Board, Geobiology, United Kingdom

Editorial Board, Aquatic Sciences, Switzerland

Editorial Board, Environmental Microbiology, Germany

Advisory Board, Stony Brook-Millstone Watershed Association

Government and Consulting

National Research Council
    Member of Panel on Marine Mineral Technolgy, 1976-1977
    Member of Committee on Ocean Waste Transportation, 1982-1984
    Member of Committee on Irrigation-Induced Water Quality Problems, 1985-1988

National Oceanic and Atmospheric Administration
    Participant in various workshops (Boulder, 1977; Seattle, 1978; Estes Park, 1978; Washington DC, 1979)
    Technical reviewer and consultant

National Advisory Committee for Oceans and Atmosphere
    Member of Panel on Ocean Mining, 1981-1982

Participant in workshops and consultant for the Environmental Protection Agency; the Bureau of Land 
    Management; the National Aeorspace Administration; and the International Joint Commission

Institut National de l'Environnement Industriel et des Risques, Membre du Conseil Scientifique (1991-present)

Nancie Centre International De L'eau, membre du Conseil d'Orientation (1995-present)

Environmental Consultant to private firms and public interest groups:
Surveyor, Nenninger & Chenevert (1972-1973); Environmental Defense Fund (1975); Nasua Corporation (1975); Water Purification Associates    (1976-1977); Massachusetts Executive Office of Environmental Affairs (1976); Procter and Gamble (1977); National Oceanic and Atmospheric Administration (1978-1979); Ocean Mining Co. (1978-1980); Pelican Pipe Co. (1980-1981); Engineering Analysis (1982); National Highway Administration (1982-1983); Perry, Dean, Stahl & Rogers, Inc. (1983); Homestake Mining Co. (1983-1991); Tetra Tech (1984-1990); MITRE Corp (1986); A.D. Little (1986); Church & Dwight (1986-present); Ciba-Geigy (1987); Alcoa (1987-present); Procter and Gamble (1987); Gradient Corp. (1987); AMOCO (1991-1995); Ropes & Gray (1994-present); Homestake Mining Co. (1997); Covington and Burling (1998); Environ Corp. (1996, 1999)


1.  Books and Special Volumes

Morel, F.M.M. and J.G. Hering, Principles and Applications of Aquatic Chemistry (2nd ed.).   John Wiley & Sons, Inc., New York, (1993).

Chisholm, S.W. and F.M.M. Morel (eds.) What regulates phytoplankton production in nutrient-rich areas of the open sea?  Limnol. & Oceanogr., Vol. 36, No. 8, (1991).

Dzombak, D.A. and F.M.M. Morel,  Surface Complexation Modeling:  Hydrous Ferric Oxide.  John Wiley & Sons, Inc., New York (1990).

Morel, F.M.M., Principles of Aquatic Chemistry.  John Wiley & Sons, Inc., New York, (1983).

2.  Book Chapters

Morel, F.M.M.  Broadening the Scientific Ph.D.:  The Princeton Experience. Proceedings of NRC's Chemical Sciences Roundtable (CSR) Workshop, Graduate Education in the Chemical Sciences:  Issues for the 21st Century, p. 130-134 (2000).

Morel, F.M.M.  Use of Trace Metals in Marine Bioremediation:  A Need for Fundamental Knowledge.  Reviews of Proceedings of the Workshop on Opportunities for Environmental Applications of Marine Biotechnology, p. 184-192 (2000).

Price, N.M. and F.M.M. Morel.  Biological cycling of iron in the sea.  In: Iron Transport and Storage in Microorganisms, Plants, and Animals, Vol. 35, p. 1-36 of 'Metal Ions in Biological Systems',  A. Sigel and H. Sigel, eds.; M. Dekker, Inc., New York, (1998).

Price, N.M. and F.M.M. Morel.  Trace metal nutrition and toxicity in phytoplankton. Acvhiv. für Hydrology.  Advances in Limnology.  42: Algae & Water Sollution.  Chap. 3 - p. 79-97.   L.C. Rai, J.P. Gaur & C.J., eds.,  Stuttgart, (1994).

Morel, F.M.M., D.A. Dzombak and N.M. Price.  Heterogeneous reactions in coastal waters.  In:  Ocean Margin Processes in Global Change, Mantoura, Martin, and Wollast (eds.), Dahlem Konferenzen.  John Wiley, New York (1991).

Hering, J. and F.M.M. Morel.  The kinetics of trace metal complexation:  Implications for metal reactivity in natural waters.  In:  Aquatic Chemical Kinetics, W. Stumm, (ed.), John Wiley, New York (1990).

Price, N.M. and F.M.M. Morel.  The role of extracellular enzymatic reactions in natural waters.  In:  Aquatic Chemical Kinetics, W. Stumm, (ed.), John Wiley, New York (1990).

Morel, F.M.M. and B. Palenik.  The aquatic chemistry of trace metals in biofilms.  In: Structure and Function of Biofilms, W.G. Characklis and P.A. Wilderer (eds.), Dahlem Konferenzen.  John Wiley, New York (1989).

Jones, G., F.M.M. Morel, C. Yentsch and E.A. Funkhouser.  A possible immunological approach to the study of plasmalemma redox enzymes in phytoplankton.  Lecture series in coastal and etuarine studies.  Springer-Verlag (1987).

Morel, F.M.M. and P.M. Gschwend.  The role of colloids in the partitioning of solutes in natural waters.  In:  Aquatic Surface Chemistry, W. Stumm (ed.), John Wiley, New York (1987).

Morel, F.M.M.  Trace metals-phytoplankton interaction:  An overview.  In: Biogeochemical Processes at the Land Sea Boundary, P. Lassere and J.M. Martin (eds.).  Elsevier, Amsterdam (1986).

Morel, F.M.M. and R.J.M. Hudson.  The geobiological cycle of trace elements in aquatic systems: Redfield Revisited.  In: Chemical Processes in Lakes, W. Stumm (ed.).  John Wiley, New York (1985).

Morel, F.M.M. and S.L. Schiff.  Geochemistry of municipal waste in coastal waters.  In:  Ocean Discharge of Municipal Wastes:  Impact on Coastal Waters, E.P. Myers (ed.), MIT Sea Grant College Program Publishers (1984).

Morel, F.M.M. and N.M.L. Morel-Laurens.  Trace Metals and plankton in the oceans:  Facts and speculations.  In:  Trace Metals in Sea Water, C.S. Wong, E. Boyle, K.W. Bruland, J.D. Burton and E.D. Goldberg (eds.), NATO Conference Series, Series IV:  Marine Chemistry, Plenum Press (pp. 841-869) (1983).

Morel, F.M.M., J.G. Yeasted and J.C. Westall  Adsorption models:  A mathematical analysis in the framework of general equilibrium calculations.  In:  Adsorption of Inorganics at the Solid-Liquid Interface, M.A. Anderson and A. Rubin (eds.), Ann Arbor Science, Ann Arbor, MI, pp.263-294 (1981).

Chisholm, S.W., F.M.M. Morel and W.S. Slocum.  The phasing and distribution of cell division cycles in marine diatoms.  In:  Primary Productivity in the Sea, P. Falkowski (ed.), Brookhaven Symposium in Biology No. 31, Plenum Press (1980).

Anderson, D.M. and F.M.M. Morel.  Toxic dinoflagellate blooms in the Cape Cod region of Massachusetts.  In: Toxic Dinoflagellate Blooms, D.L. Taylor and H.H. Seliger (eds.), Proceedings of the Second  International Conference on Toxic Dinoflagellate Blooms, Key Biscayne, FL (1979).

Morel, F.M.M., N.M.L. Morel-Laurens, D.M. Anderson, D.M. McKnight and J.G. Reuter, Jr., Trace metal speciation and toxicity in phytoplankton cultures.  In:  Advances in Marine Environmental Research.  Proceedings of a Symposium, EPA-600/9-79-035 (1979).

Morel, F.M.M. and J.G. Yeasted.  On the interfacing of chemical, physical and biological water quality models.  In:  Fate of Pollutants in the Air and Wave Environment, I.H. Suffet (ed.), Wiley-Interscience (1977).

Morel, F.M.M., R.E. McDuff and J.J. Morgan.  Interactions and chemostasis in aquatic chemical systems.  In:  Trace Metals and Metal-Organic Interactions in Natural Waters, P.C. Singer (ed.), Ann Arbor Science, Ann Arbor, MI (1973).

3.  Journal Articles

Ekstrom, E.B., F.M.M. Morel, J.M. Benoit. Mercury Methylation Independent of Acetyl-CoA Pathway in SRB. Appl. and Environ. Microbiol (submitted).

Kraepiel, A.M.L., K. Keller, H.B. Chin, E.G. Malcolm, F.M.M. Morel, Sources and Variations of Mercury in Tuna Environ. Sci. Technol. (submitted).

Ho, T.-Y., A. Quigg, Z.V. Finkel, A.J. Milligan, K. Wyman, P.G. Falkowski, F.M.M. Morel

Saito, M. A., D.M. Sigman and F.M.M. Morel. The Bioinorganic Chemistry of the Ancient Ocean: The Co-Evolution of Cyanobacterial Metal Requirements and Biogeochemical Cycles at the Archean-Proterozoic Boundary. Inorganica Chimica Acta (in press).

Milligan, A. J. and F.M.M. Morel. Dynamics of silicon metabolism and silicon isotopic discrimination in a marine diatom as a function of pCO2 Limnol. Oceanogr. (in press) .

Morel, F.M.M. and N.M. Price, The Biogeochemical Cycles of Trace Metals in the Oceans. Science, 300: 944-947 (2003).

Newman, D.K., R.S. Oremland, P.R. Dowdle, F.M.M. Morel, and J.F. Stolz.  Purification and characterization of the dissimilatory arsenate reductase from Sulfurospirillum barnesii strain SES-3.  Proc. Nat. Acad. Sci., (in revision).

Milligan, A.J. and F.M.M. Morel.  A proton buffering role for silica in diatoms.  Science, 297: 1848-1850 (2002).

Tortell, P.D., G.R. DiTullio, D.M. Sigman, and F.M.M. Morel.  CO2 effects on taxonomic composition and nutrient utilization in an Equatorial Pacific phytoplankton assemblage. MEPS, 236: 37-43 (2002).

Tortell, P.D. and F.M.M. Morel.  Sources of inorganic carbon for phytoplankton in the Eastern Subtropical and Equatorial Pacific Ocean.  Limnol. Oceanogr., 47: 1012-1022 (2002).

Morel, F.M.M., E.H. Cox, A.M.L. Kraepiel, T.W. Lane, A.J. Milligan, I. Schaperdoth, J.R. Reinfelder and
P.D. Tortell.  Acquisition of inorganic carbon by the marine diatom Thalassiosira weissflogii. Functional Plant
Biology, 29: No.'s 2 & 3, pp. 301-308 (2002).

Gawel, J.E., C.G. Trick, and F.M.M. Morel.  Phytochelatins are bioindicators of atmospheric metal insult to trees near Sudbury, Ontario, Canada.  Environ. Sci. Tech., 35: 2108-2113 (2001).

Lam, P.J., P.D. Tortell, and F.M.M. Morel.  Iron additions decrease the ratio of inorganic to organic carbon production in the ocean.  Limnol. Oceanogr., 46: 1199-1202 (2001).

Lalonde, J.D., M. Amyot, A.M.L. Kraepiel, and F.M.M. Morel.  Photooxidation of Hg(0) in an estuarine system.  Environ. Sci. Technol.,
35: 1367-1372 (2001).

Jay, J., F.M.M. Morel, and H.F. Hemond.  Mercury speciation in the presence of polysulfides. Environ. Sci. Tech.,
34: 2196-2200 (2000).

Reinfelder, J.R., A.M.L. Kraepiel, and F.M.M. Morel.  Unicellular C4 photosynthesis in a marine diatom.  Nature, 407: 996-999 (2000).

Keller, K., K. Tan, F.M.M. Morel, and D.F. Bradford.  The North Atlantic thermohaline circulation collapse as a constraint on economically optimal carbon dioxide emissions. Working Paper Series 7476, NBER (2000).  Climatic Change, 47: 17-43 (2000).

Cox, E.H., G.L. McLendon, F.M.M. Morel, T.W. Lane, R.C. Prince, I.J. Pickering, and G.N. George.  The active site structure of Thalassiosira weissflogii carbonic anhydrase 1.  Biochemistry, 39: 12128-12130 (2000).

Riebesell, U., I. Zondervan, B. Rost, P.D. Tortell, R.E. Zeebe, and F.M.M. Morel.  Reduced calcification in marine plankton in response to increased atmospheric CO2.  Nature, 407: 364-367 (2000).

Tortell, P.D., G.H. Rau, and F.M.M. Morel.  Inorganic carbon acquisition in coastal Pacific phytoplankton communities.  Limnol. Oceanogr., 45: 1485-1500 (2000).

Lane, T.W. and F.M.M. Morel.  Regulation of carbonic anhydrase expression by zinc, cobalt, and carbon dioxide in the marine diatom Thalassiosira weissflogiiPlant Physiology, 123: 345-352 (2000).

Lane, T.W. and F.M.M. Morel.  A biological function for cadmium in marine diatoms.  PNAS, 97: 4627-4631 (2000).

Ahner, B. and F.M.M. Morel.  Phytochelatins in microalgae.  Progress in Phycological  Research,13: 1-31 (1999).

Cullen, J.T., T.W. Lane, F.M.M. Morel, and R.M. Sherrell.  Modulation of cadmium uptake in phytoplankton by seawater CO2 concentration.  Nature, 402: 165-167 (1999).

Keller, K. and F.M.M. Morel.  A model of carbon isotopic fractionation and active carbon uptake in phytoplankton. Marine Ecology Progress Series, 182: 295-298 (1999).

Kraepiel, A.M.L., K. Keller, and F.M.M. Morel.  A model for metal adsorption on clays.  J.  Colloid and Interface Science,210: 43-54 (1999).

Ahner, B.A., J.G. Lee, N.M. Price, and F.M.M. Morel.  Phytochelatin concentrations in the equatorial Pacific.  Deep-Sea Res., 45:1779-1796 (1998).

Morel, F.M.M., A.M.L. Kraepiel, and M. Amyot.  The chemical cycle and bioaccumulation of mercury. Annual Review Ecology and Systematics, Vol. 29:  543-566 (1998).

Gueneau, P., F.M.M. Morel, J. Laroche, and D. Erdner.  The PetF region of the cholorplast genome from the diatom Thalassiosira weissflogii,  sequence, organization and phylogeny.  European Journal of Phycology, 33:  203-211 (1998).

Kraepiel, A.M.L, K. Keller, and F.M.M. Morel.  On the acid-base chemistry of permanently charged minerals.  Envi. Sci. Tech., 32: 2829-2838 (1998).

Newman, D.K., D. Ahmann, and F.M.M. Morel.  A brief review of dissimilatory arsenate reduction. Geomicrobio. Journ., 15: 255-268 (1998).

Morel, F.M.M. and A. Kraepiel.  Coulombic effects on the adsorption of trace cations on clays.  J. Cont. Hydrology, 28: 17-20 (1997).

Morel, F.M.M. Discussion on "A mechanistic description of Ni and Zn sorption on Namontmorillonite.  Part I: Titration and sorption measurements.  Part II: Modeling" by Bart Baeyens and Michael H. Bradbury.  J. Cont. Hydrology, 28: 7-10 (1997).

Amyot, M., G.A. Gill, and F.M.M. Morel.  Production and loss of dissolved gaseous mercury in coastal seawater.  Env. Sci. Tech., 31: 3606-3611 (1997).

Tortell, P.D., J.R. Reinfelder, and F.M.M. Morel.  Active uptake of bicarbonate by diatoms. Nature, 390: 243-244 (1997).

Roberts, S.B., T. Lane, and F.M.M. Morel.  Carbonic anhydrase in the marine diatom Thalassiosira weissflogii (bacillariophyta).  J. Phycol., 33: 845-850 (1997).

Ahmann, D., L.R. Krumholz, H.H. Hemond, D.R. Lovley, and F.M.M. Morel.  Microbial mobilization of arsenic from sediments of the Aberjona Watershed. Env. Sci. Tech.,31: 2923-2930 (1997).

Newman, D.K., E.K. Kennedy, J.D. Coates, D. Ahmann, D.J. Ellis, D.R. Lovley, and F.M.M. Morel.  Dissimilatory As(V) and S(VI) reduction in Desulfotomaculum aurpigmentum, sp. nov. Archives Microbiol., 168: 380-388 (1997).

Ahner, B.A., F.M.M. Morel, and J.W. Moffett.  Trace metal control of phytochelatin production in coastal waters.  Limnol. Oceanogr., 42: 601-608 (1997).

Newman, D.K., T.J. Beveridge and F.M.M. Morel.  Precipitation of As2S3 by a novel As(V)- reducing bacterium, strain OREX-4.  Appl. Env. Microbiol., 63: 2022-2028 (1997).

Kraepiel, A.M.L., J.-F. Chiffoleau, J.-M. Martin, and F.M.M. Morel.  Geochemistry of trace metals in the Girone estuary.  Geochim. Cosmochim. Acta,  61: 1421-1435 (1997).

Voelker, B., F.M.M. Morel, and B. Sulzberger.  Iron redox cuycling in surface waters: effects of humic substances and light.  Env. Sci. Tech., 31: 1004-1011 (1997).

Gawel, J.E., B.A. Ahner, A.J. Friedland and F.M.M. Morel.  Role for heavy metals in forest decline Indicated by phytochelatin measurements. Nature, 381:  64-65 (1996).

Lee, J.G., B.A. Ahner and F.M.M. Morel.  Export of cadmium and phytochelatin by the marine diatom Thalassiosira weissflogiiEnv. Sci. Tech., 30: 1814-1821 (1996).

Mason, R.P., J.R. Reinfelder and F.M.M. Morel.  The uptake, toxicity, and trophic transfer of mercury in a coastal diatom.  Env. Sci. Tech., 30:  1835-1845 (1996).

Yee, D. and F.M.M. Morel.  In vivo substitution of zinc by cobalt in carbonic anhydrase of a marine Diatom.  Limnol. Oceanogr., 41: 573-577 (1996).

Joseph, E.M. and F.M.M. Morel and N.M. Price.  Effect of aluminum on surfase phosphateses of Chlamydomonas reinhardtii.  Can. J. Fish. Aquat. Sci., 52: 353-357 (1995).

Lee, J.G. and F.M.M. Morel.  Replacement of zinc by cadmium in marine phytoplankton. Mar. Ecol. Prog. Ser., 127: 305-309 (1995).

Lee, J.G., S.R. Roberts and F.M.M. Morel.  Cadmium: a nutrient for the marine diatom Thalassiosira weissflogii.  Limnol. & Oceanogr., 40: 1056-1063 (1995).

Mason, R.P., F.M.M. Morel and H.F. Hemond.  The role of microorganisms in elemental mercury formation in natural waters.  Water Air Soil Pollut., 80:  775-787 (1995).

Mason, R.P., J.R. Reinfelder, and F.M.M. Morel.  Bioaccumulation of mercury and methylmercury. Water Air Soil Pollut., 80: 915-921 (1995).

Ahner, B.A., S. Kong and F.M.M. Morel.  Phytochelatin synthesis in marine algae: I. An interspecies comparison.  Limnol. & Oceanography, 40(4): 649-657 (1995).

Ahner, B.A. and F.M.M. Morel.  Phytochelatin production in marine algae: II. Induction by various metals. Limnol & Oceanography, 40(4) 658-665 (1995).

Ahmann, D., A.L. Roberts, L.R. Krumholz, and F.M.M. Morel.  Microbe grows by reducing arsenic. Nature, 371: 750 (1994).

Ahner, B.A., N.M. Price, and F.M.M. Morel.  Photochelatin production by marine phytoplankton and low free metal ion concentrations:  Lab studies and field data for Massachusetts Bay.  Proc. Natl. Acad. Sci., 91:  8433-8436 (1994).

Morel, F.M.M., J.R. Reinfelder, S.B. Roberts, C.P. Chamberlain, J.G. Lee, and D. Yee.  Zinc and carbon co-limitation of marine phytoplankton. Nature, 369: 740-742 (1994).

Mason, R.P., W.F. Fitzgerald and F.M.M. Morel.  The biogeochemical cycling of elemental mercury:  Anthropogenic Influences.  Geochim. Cosmochim. Acta, 58: 3191-3198 (1994).

Price, N.M., B.A. Ahner and F.M.M. Morel.  The Equatorial Pacific Ocean:  grazer-controlled phytoplankton populations in iron-limited ecosystem. Limnol. & Oceanogr., 39: 520-534 (1994).

Hudson, R.J.M. and F.M.M. Morel.  Trace metal transport by marine microorganisms: implications of metal coordination kinetics.  Deep-Sea Res., 40: 129-150 (1993).

Mason, R.P. F.M.M. Morel, and H.F. Hemond.  The role of microorganisms in elemental mercury formation in natural waters.  In:  Heavy Metals in the Environment (J.O. Nriagu and R.P. Allan, eds.) Vol. 2 pp. 293-296 (1993).

Hudson, R.J.M., D.T. Covault and F.M.M. Morel.  Investigations of iron coordination and redox reactions in seawater using 59Fe radiometry and ion-pair solvent extraction of amphiphilic iron complexes.  Marine Chem., 38: 209-235 (1992).

Bartschat, B.M., S.E. Cabaniss and F.M.M. Morel.  An oligoelectrolyte model for cation binding by humic substances.  Env. Sci. Tech., 26: 284-294 (1992).

Green, S.A., F.M.M. Morel and N.V. Blough.  Investigation of the electrostatic properties of humic substances by fluorescence quenching.  Env. Sci. Tech., 26: 294-302 (1992).

Palenik, B., D.J. Kieber and F.M.M. Morel.  Dissolved organic nitrogen use by phytoplankton:  The role of cell-surface enzymes.  Biological Oceanography, 6: 347-354 (1991).

Morel, F.M.M., R.J.M. Hudson and N.M. Price.  Limitation of productivity by trace metals in the sea.  Limnol. & Oceanogr., 36: 1742-1755 (1991).

Price, N.M. and F.M.M. Morel.  Co-limitation of phytoplankton growth by nickel and nitrogen.  Limnol. & Oceanogr., 36: 1071-1077 (1991).

Morel, F.M.M., J.G. Rueter, and N.M. Price.  Iron nutrition of phytoplankton and its possible importance in the ecology of ocean regions with high nutrient and low biomass. Oceanography, 4:  56-61 (1991).

Palenik, B. and F.M.M. Morel.  Amine oxidases of marine phytoplankton. Appl. Env. Microbiol., 57: 2440-2443 (1991).

Palenik, B., N.M. Price and F.M.M. Morel.  Potential effects of UV-B on the chemical environment of marine organisms: A review.  Environmen. Pollut., 70: 117-130 (1991).

Price, N.M., L.F. Andersen and F.M.M. Morel.  Iron and nitrogen nutrition of equatorial Pacific plankton. Deep-Sea Res., 38: 1361-1378 (1991).

Hering, J.G. and F.M.M. Morel.  Kinetics of trace metal complexation: Ligand exchange reactions.  Env. Sci. Tech., 24: 242-252 (1990).

Hudson, R.J.M. and F.M.M. Morel.  Iron transport in marine phytoplankton:  Kinetics of cellular and and medium coordination reactions.  Limnol. & Oceanogr., 35: 998-1016 (1990).

Newman, K.A., F.M.M. Morel and K.D. Stozenbach.  Settling and coagulation characteristics of fluorescent particles determined by flow cytometry and fluorometry. Env. Sci. Tech., 24: 506-513 (1990).

Nirel, P.M. and F.M.M. Morel.  Comment: Pitfalls of sequential extractions. Water Research, 24: 1055-1056 (1990).

Palenik, F. and F.M.M. Morel.  Amino acid utilization by marine phytoplankton:  A novel mechanism.   Limnol. & Oceanogr., 35: 260-269 (1990).

Palenik, B. and F.M.M. Morel.  Comparison of cell-surface L-amino acid oxidases from several marine phytoplankton. Mar. Ecol. Prog. Ser., 59:  195-201 (1990).

Price, N.M. and F.M.M. Morel.  Cadmium and cobalt substitution for zinc in a zinc-deficient marine diatom. Nature, 344: 658-660 (1990).

Rich, H.W. and F.M.M. Morel.  Availability of well-defined iron colloids to the marine diatom Thalassiosira weissflogiiLimnol. & Oceanogr., 35:  652-662 (1990).

Roulier, M.A., B. Palenik and F.M.M. Morel.  A method for the measurement of choline and hydrogen peroxide in seawater.  Marine Chem., 30: 409-421 (1990).

Cabaniss, S.E. and F.M.M. Morel.  Comment on "A unified physicochemical description of the protonation and metal ion complexation equilibrium of natural organic acids, Part 1 & 2."  Env. Sci. Tech., 23:  746-747 (1989).

Hering, J.G. and F.M.M. Morel.  Slow coordination reactions in seawater. Geochim. Comsochim. Acta, 53: 611-618 (1989).

Hudson, R.J.M. and F.M.M. Morel.  Distinguishing between extra- and intracellular iron in marine phytoplankton.  Limnol. & Oceanogr., 34: 1113-1120 (1989).

Price, N.M., G.I. Harrison, J.G. Hering, R.J. Hudson, P.M.V. Nirel, B. Palenik and F.M.M. Morel.  Preparation and chemistry of the artificial algal culture medium aquil. Biolog. Oceanogr., 6: 443-461 (1988/1989).

Hering, J.G. and F.M.M. Morel.  Kinetics of trace metal complexation:  Role of alkaline earth metals. Env. Sci. Tech., 22: 1469-1478 (1988).

Palenik, B. and F.M.M. Morel.  Dark production of H2O2 in the Sargasso Sea.  Limnol. & Oceanogr., 33 (6:2)  1606-1611 (1988).

Hering, J.G. and F.M.M. Morel.  Humic acid complexation of calcium and copper. Env. Sci. Tech., 22: 1234-1237 (1988).

Morel, F.M.M.  Solute-particle and particle-particle rections in seawater. Appl. Geochem. 3:80 (1988).

Jones, G.J. and F.M.M. Morel.  Plasmalemma redox activity in the diatom Thalassiosira. Plant Physiol., 87: 143-147 (1988).

Palenik, B., F.M.M. Morel and O.C. Zafiriou.  Hydrogen peroxide production by a marine phytoplankton. Limnol. & Oceanogr., 32(6): 1365-1369 (1987).

Hering, J.G., F.M.M. Morel, W.G. Sunda and R.L. Ferguson.  A field comparison of two methods for the determination of copper complexation:  Bacterial bioassay and fixed-potential aperometry.  Marine Chemistry, 20: 255-312 (1987).

Dzombak, D.A. and F.M.M. Morel.  Adsorption of inorganic pollutants in aquatic systems. J. Hydraulic Eng.  ASCE. 113: 430-475 (1987).

Dzombak, D.A. and F.M.M. Morel.  Development of a data base for modelling adsorption of inorganics on iron and aluminum oxides.  Environmental Progress AICHE 6: 133-137 (1987).

Morel, F.M.M.  Kinetics of nutrient uptake and growth in phytoplankton.  J. Phycol., 27: 137-150 (1987).

Jones, G.J., B. Palenik and F.M.M. Morel.  Trace metal reduction by phytoplankton:  The role of plasmalemma redox enzymes.  J. Phycol., 23: 237-244 (1987).

Coudrain-Ribstein, A. and F.M.M. Morel.  Transport modeling of major reactive elements in a variable temperature field:  Methodology and simple examples.  Bulletin de la Societe Geologique de France  Numero Special 5:  1009-1017 (1987).

Cherry, J.A., F.M.M. Morel, J.V. Rouse, J.L. Schnoor and M.G. Wolman.  Hydrogeochemistry of sulfide and arsenic-rich tailings and alluvium along Whitewood Creek, South Dakota.  Three part series.  Mineral and Energy Resources, Colorado School of Mines (1986).

Harrison, G.I. and F.M.M. Morel.  Response of the marine diatom Thalassiosira weissflogii to iron stress. Limnol. & Oceanogr., 31 (5): 989-997 (1986).

Dzombak, D.A. and F.M.M. Morel.  Sorption of cadmium on hydrous ferric oxide at high sorbate/sorbent ratios:  Equilibrium and modeling.  J. Coll. & Inter. Sci., 112(2): 588-598 (1986).

Dzombak, D.A., W. Fish and F.M.M. Morel.  Metal-humate interactions, 1:  Discrete ligand and continuous distribution models.  Env. Sci. Tech., 20(7): 669-675 (1986).

Fish, W., D.A. Dzombak and F.M.M. Morel.  Metal-humate interactions, 2:  Applications and comparison of models.  Env. Sci. Tech., 20(7): 676-683 (1986).

Farley, K. and F.M.M. Morel.  The role of coagulation in the kinetics of sedimentation. Env. Sci. Tech., 20(2): 187-195 (1986).

Watras, C.J. and F.M.M. Morel.  Nickel accumulation by Scenedesmus and Daphnia:  Food-chain transport and geochemical implications.  Can. J. Fish. Aquat. Sci., 42(4): 724-730 (1985).

Fish, W. and F.M.M. Morel.  Propagation of error in fulvic acid titration data:  A comparison of three analytical methods.  Can. J. Chem., 63(6): 1185-1193 (1985).

Farley, K.J., D.A. Dzombak and F.M.M. Morel.  A surface precipitation model for the sorption of cations on metal oxides.  J. Coll. Inter. Sci., 106:1 (1985).

Waite, T.D. and F.M.M. Morel.  Photoreductive dissolution of colloidal iron oxides in natural waters. Env. Sci. Tech., 18: 860-868 (1984).

Waite, T.D. and F.M.M. Morel, Ligand exchange and fluorescence quenching studies of the fulvic acid-iron interaction:  Effects of pH and light. Anal. Chemica Acta, 162: 263-264 (1984).

Waite, T.D. and F.M.M. Morel.  Photoreductive dissolution of colloidal iron oxide:  Effect of citrate. J. Coll. Inter. Sci., 102: 1 (1984).

Waite, T.D. and F.M.M. Morel.  Coulometric technique for studying the redox dynamics of iron in seawater. Anal. Chem., 56: 787-792 (1984).

Harrison, G.I. and F.M.M. Morel.  Antagonism between cadmium and iron in the marine diatom Thalassiosira weissflogiiJ. Phycol., 19: 495-507 (1983).

Fish, W. and F.M.M. Morel.  Characterization of organic copper-complexing agents released by Daphnia magna.  Can. J. Fish. Aquatic Sci., 40: 1270 (1983).

Waite, T.D. and F.M.M. Morel.  Characterization of complexing agents in natural waters by Cu(II)/Cu(I) amperometry.  Anal. Chem., 55: 1268-1274 (1983).

Anderson, M.A. and F.M.M. Morel.  The influence of aqueous iron chemistry on the uptake of iron by the coastal diatom Thalassiosira weissflogii. Limnol. & Oceanogr., 27: 745-752 (1982).

Foster, P.L. and F.M.M. Morel.  Reversal of cadmium toxicity in the diatom Thalassiosira weissflogii.Limnol. & Oceanogr., 27: 745-752 (1982).

Reuter, J.G., Jr. and F.M.M. Morel.  Interactions between zinc deficiency and copper toxicity as it affects the silicic acid uptake in the marine diatom, Thalassiosira pseudonanaLimnol. & Oceanogr., 26: 67-73 (1981).

Rueter, J.G., Jr., S.W. Chisholm and F.M.M. Morel.  The effect of copper toxicity on silicic acid uptake and growth in Thalassiosira pseudonana (Bacillariophyceae). J. Phycol., 17: 270-278 (1981).

McDuff, R.E. and F.M.M. Morel.  Geochemical control of seawater alkalinity (Sillén revisited).  Env. Sci. Tech., 14:1182 (1980).

Anderson, M.A. and F.M.M. Morel.  Uptake of Fe(II) by a diatom in oxic culture medium. Mar. Bio. Let., 1:263 (1980).

Swallow, K.C., D.N. Hume and F.M.M. Morel.  Sorption of copper and lead by hydrous ferric oxide. Env. Sci. Tech., 14:1326 (1980).

McKnight, D.M. and F.M.M. Morel.  Copper complexation by siderophores from filamentous blue-green algae.  Limnol. & Oceanogr., 25:62 (1980).

Westall, J.C., F.M.M. Morel and D.N. Hume.  Chloride interference in cupric ion selective electrode measurements.  Anal. Chem., 51:1792 (1979).

McKnight, D.M. and F.M.M. Morel.  Release of weak and strong copper complexing agents by algae. Limnol. & Oceanogr. 24(5):823-837 (1979).

Morel, F.M.M., J.G. Rueter, Jr., D.M. Anderson and R.R.L. Guillard.  AQUIL:  A chemical defined phytoplankton culture medium for trace metal studies. J. Phycol., 15:135 (1979).

Anderson, D.M. and F.M.M. Morel.  The seeding of two red tide blooms by the germination of benthic Gonyaulax tamarensis Hypnocysts.  J. Est. and Coast. Mar. Sci., 8:279 (1979).

Anderson, M.A., F.M.M. Morel and R.R. L. Guillard.  Growth limitation of a coastal diatom by low zinc ion activity.  Nature, 276(5683): 70-71 (1978).

Yeasted, J.G. and F.M.M. Morel.  Empirical insights into lake response to nutrient loadings with application to models of phosphorus in lakes.  Envir. Sci. Tech., 12:195 (1978).

Morel, N.M.L., J.G. Rueter, Jr. and F.M.M. Morel.  Copper toxicity to Skeletonema costatum (Bacillariophyceae). J. Phycol., 14:43 (1978).

Swallow, K.C., D.M McNight, J.C. Westall, N.M.L. Morel and F.M.M. Morel.  Potentiometric determiniation of coper complexation by phytoplankton exudates. (Note).  Limnol. & Oceanogr., 23:538 (1978).

Anderson, D.M. and F.M.M. Morel.  Copper sensitivity of Gonyaulax tamarensis. Limnol & Oceanogr., 23: 296 (1978).

Morel, F.M.M., R.E. McDuff and J.J. Morgan.  Thoery of interaction intensities, buffer capacities and pH stability in aqueous systems, with application to the pH of seawater and a heterogeneous model ocean system.  J. Mar. Chem., 4:1 (1976).

Morel, F.M.M., J.C. Westall, C.R. O'Melia and J.J. Morgan.  Fate of trace metals in the Los Angeles County wastewater discharge.  Env. Sci. Tech., 9:256 (1975).

Morel, F.M.M.  A study of passive potassium efflux from human red blood cells using iron specific electrodes.  J. Membr. Bio., 12:68 (1972).

Morel, F.M.M. and J.J. Morgan.  A numerical method for computing equilibria in aqueous chemical systems. Envir. Sci. and Tech., 6:58 (1972).

Morel, F.M.M., R.F. Baker and H. Wayland.  Quantitation of human red blood cell fixation by glutaraldehyde.  J. Cell Biol., 48:91 (1971).

4.  Principal Technical Reports and Technical Notes
TR 259  Morel, F.M.M. and S.L. Schiff.  Geochemistry of municipal waste in coastal waters.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA (1980).

Contributing author to Dahlem Konferenzen report on "Organic Acids in Natural Waters," John Wiley, New York (1989).

Contributing author to "Scientific Problems Relating to Ocean Pollution."  E.D. Goldbert (ed.), Proceedings of a Workshop, Estes Park, CO, July 1978, NOAA/ERL (1979).

Contributing author to "Trace Metals and Marine Production Processes," R.A. Feely and H. Curl, Jr. (eds.), NOAA Special Report, NOAA/ERL #385 (1978).

TN 19  Westall, J.C. and F.M.M. Morel.  FITEQL: A general algorithm for determination of metal-liqand complex stability constants from experimental data.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA  (1977).

TN 18  Westall, J.C., J.L. Zachary and F.M.M. Morel.  MINEQL:  A compact program for computation of chemical equilibria in aquatic systems.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA (1976).

TN 16  Morel, F.M.M., J.C. Westall, J.G. Rueter, Jr. and J.P. Chaplick.  Description of algal growth media AQUIL and FRAQUIL.  R.M. Parsons Laboratory for Water Resources and Hydrodynamics, Massachusetts Institute of Technology, Cambridge, MA (1975).

EQ,73-02.  McDuff, R.E. and F.M.M. Morel.  Description of the chemical equilibrium program REDEQL2.  Keck Laboratory of Environmental Engineering Science, California Institute of Technology, Pasadena, CA (1973).

EQ.72-01.  McDuff, R.E. and F.M.M. Morel.  REDEQL, a general program for the computation of chemical equilibrium in aqueous systems.  Keck Laboratory of Environmental Engineering Science, California Institute of Technology, Pasadena, CA (1972).

Former Doctoral Advisees and Postdoctoral Associates

Philippe Tortell (Ph.D., 2001) Assistant Professor, University of British Columbia, Canada

Klaus Keller (Ph.D., 2000) Assistant Professor, Penn State University, PA

Anne Kraepiel (Ph.D., 1999) Assistant Professor, Centre de Geochimie de Surface, Strasbourg, France

Jenny Jay, (Ph.D., 1999) Postdoctoral Fellow, Harvard University, MA

Pulcherie Gueneau (Ph.D., 1998) Assistant Professor, Simon Bolivar University, Caracas, Venezuela

Diane Newman (Ph.D., 1997) Assistant Professor - Division of Geological and Planetary Sciences, CalTech

James Gawel (Ph.D., 1996) Assistant Professor, University of Washington, WA

Donald Yee (Ph.D., 1996) Assistant Environmental Scientist, San Francisco Estuary Institute, CA

Dianne Ahmann (Ph.D., 1996) Assistant Professor, Colorado School of Mines, CO

John Reinfelder (Post-doc., 1992-1996) Assistant Professor, Rutgers University

Jennifer G. Lee (Ph.D., 1995) Research Associate, University of Maryland, MD

Bettina (Bartschat) Voelker (Ph.D., 1994 Zurich) Assistant Professor, MIT, MA

Beth A. Ahner (Ph.D., 1994) Assistant Professor, Cornell University, Ithaca, NY

Sarah Green (Ph.D., 1992) Assistant Professor, Michigan Tech, MI

Neil M. Price (Post-doc., 1987-1991) Professor, McGill University, Montreal, Canada

Brian Palenik (Ph.D., 1989) Associate, Professor, Scripps Institution of Oceanography, UCSD, CA

Robert Hudson (Ph.D., 1989) Assistant Professor, University of Illinois, IL

Janet Hering (Ph.D., 1988) Professor, Caltech, CA

Gary Jones (Post-doc., 1986-1988) Div. Of Water Res., CSIRO, Griffith, NSW 2680, Australia

David Dzombak (Ph.D., 1986) Professor, Carnegie Mellon University, PA

Kevin Farley (Ph.D., 1984) Professor, Manhattan College, NY

William Fish (Ph.D., 1984) Associate Professor, Oregon Graduate Institute, Beaverton, OR

T. David Waite (Ph.D., 1984)  Professor and Chairman, School of Civil Engineering, University of New South Wales, Australia

Carl Watras (Post-doc., 1981-1983) Research Assoc., Univ. of Wisconsin, Boulder Junction, WI

Patricia Foster (Post-doc. 1980-1982) Associate Professor of Env. Health, School of Public Health, Boston University, MA

John Rueter (Ph.D., 1979) Professor of Biology and Director of Env. Sci. Program, Portland State University, OR

Donald Anderson (Ph.D., 1978) Senior Scientist, Woods Hole Oceanographic Institution, MA

Diane McKnight (Ph.D., 1978) Professor, University of Colorado, Boulder, CO

Joseph Yeasted (Ph.D., 1978) Director of Major Projects, Int. Tech. Corp., Green Valley, NV

David Burmaster (Ph.D., 1978) President, Alceon Corporation, Cambridge MA

Kathleen Swallow (Ph.D., 1978) Professor, Merrimack College, North Andover, MA

John Westall (Ph.D., 1978) Professor, Dept. of Chemistry, Oregon State University, OR

Harry Hemond (Ph.D. 1977) Professor and Director, Parsons Lab., MIT, MA

Updated 9/10/03